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ages to produce—great and sudden physical modifications might often
produce the extinction of a race just as it was approaching perfection,
and a hundred checks, of which we can know nothing, may have re
tarded the progress towards perfect adaptation ; so that we can hardly
wonder at there being so few cases in which a completely successful
result has been attained, as shown by the abundance and wide
diffusion of the creatures so protected.”—(Contributions. ftp. 68-69.)
Whichever of these two authorities we follow, defensive
colouring cannot be said to afford the most favourable
illustration of the action of Natural Selection. If Mr.
Bates’ view is adopted, the part played is very slight, and
one does not see the necessity for the introduction of
Natural Selection for which he contends. If Mr. Wallace’s
view be correct, then this department does not offer a very
favourable illustration of the action of Natural Selection,
seeing that the process is so difficult and so protracted and
so often accompanied by failure.
But in this particular case, we are not at liberty to leave
the actual amount of difference an open question and to
argue from either hypothesis as to the possible result. For
in the case of defensive colouring the initial change must
be considerable before Natural Selection can come into
action. There must be enough likeness to deceive before
the individual so favoured can be saved.
But, for the sake of argument, let us take the two
hypotheses of slight changes and of considerable changes
in each generation, and let us see what would be the effect
upon the supposed action of Natural Selection. The
agent in this case is the bird or beast of prey which acts
efficiently by virtue of its keenness of vision, and it is one
of the commonplaces of the theory that the vision of the
enemy is quickened by the protracted process. Let us see
what the result must be, first, in the case of considerable
variations. If the process of Natural Selection begins at
that point where certain insects pass into invisibility, or are