5
variations in fishes, whilst living in their natural state, and distinct
races have not been formed ; on the other hand, a closely allied
species—the gold-fish—from being reared in glass or open vessels,
and from having been carefully attended to by the Chinese, has
yielded many races.”—(The Variation of Animals and Plants under
Domestication, vol. ii.,fi. 236.)
Without selection individual differences will be swallowed
up in the average of the race, through the principle which
Mr. Francis Galton calls the regression to mediocrity.
This principle differs from the ordinary action of atavism :
which, as I have pointed out, will sometimes cause an
individual to resemble a remote ancestor, and which pro
duces an influence not to be easily calculated. The
regression to mediocrity, on the other hand, represents the
effect of the ancestry as a whole ; and Mr. Galton seems to
have succeeded in ascertaining the exact amount of this
influence. This principle, I venture to believe, only applies
to those variations which are inevitably and necessarily
associated with sexual reproduction. Thus understood, it
involves the absolute necessity for the strictest isolation
of similar variants, but at the same time, it cannot be
understood to mean that this regression to mediocrity will
take place when similar variants are isolated for breeding
purposes, for such an assertion would be diametrically
opposed to the experiences of the cattle breeder and the
pigeon fancier.
“ It will be seen” (says Mr. Galton) “from the large values of the
ratios of regression, how speedily all peculiarities that are possessed
by any single individual to an exceptional extent, and which blend
freely together with those of his or her spouse, tend to disappear. A
breed of exceptional animals, rigorously selected and carefully
isolated from admixture with others of the same race, would become
shattered by even a brief period of opportunity to marry freely.”—
{Nattire. vol. xxxiii.,f. 297.)
But Mr. Galton emphatically states that the law of
regression does not invalidate the principle of selection :—