525
The first argument which I propose to consider is that
drawn from the geographical distribution of animals and
plants, and from the relation which sometimes exists
between the past and the present inhabitants of any given
locality. From these phenomena it is inferred that exist
ing species “ came into existence where they now occur
by way of genetic descent, with perpetual migration and
correlative modification,” and that “ every species has
come into existence coincident both in space and time
with a pre-existing and closely-allied species.” In short,
the argument is an argument to show that the transmu
tation of species has taken place in the past even as it
is now taking place in the present.
The change of conditions which requires a readjustment
of the organism to adapt itself may arise either through
the transference of a species to other spheres, or through
the change of conditions which may occur in the same
locality from time to time. In connection with the former
problem, we have especially to consider the circumstances
under which the distribution has taken place, whether of
conscious emigrants or of the passive victims of certain
means of dispersal. We have now to consider how far this
geographical distribution has been favourable to the trans
mutation of species by means of Natural Selection.
Assuming that a species started from a given locality,
it would naturally spread in those directions where there
was an opening, and move on those lines in which there was
an unbroken and continuous connection. The principal
method of distribution would therefore be by a process of
emigration whenever a favourable outlet could be found.
The argument for Organic Evolution, drawn from geo
graphical distribution, is based upon the fact that along
such lines of unbroken communication we find a series of
allied species adapted to their different conditions. There