LAI rinn*, profile can be not correlated. Thus, if crop production results essentially from the absorbed
photosynthe.fically active radiation by a LAI “big- leaf’ in most of the production models, they are in fact, more
successful for the crops without vegetative period (winter dormancy when tillering occurs) like: maize,
sugarbeets, spring wheat... For the winter wheat, which can show important differences in the canopy
development regarding to the farming practices. Porter (1984) and his colleagues found that it was a fundamental
point to simulate correctly the shoot population and the growth of the leaves at the individual level in oder to
have a good description of the variables relevant for the yield estimation.
Because radiosity approach seems to be possible due to the architecture description made by
AFRCWHEAT, we explain here the main considerations used in the canopy development submodel. Driving
variables of the canopy development part are the daily average temperature, the time, and the daylength.
Leaf appearance: The rule used came from ground observations showing an empirical relation between the rate
of leaf appearance per degree day and the rate of change of daylength at emergence. So, in a practical way, the
phyllochron interval (thermal time required between the appearance of successive leaves), computed at the day of
emergence, is bigger for early sowing fields than for late sowed fields, because emergence day is closer to the
equinoxe date in the first case than in the second case. A mean value of the phyllochron interval is 100 - 110
degres.days.
Tiller production and survival : When the stage “4 leaves an the main shoot” is reached, tillers are produced until
the double-ridge stage. A time step of 7 days is used for die simulation: the number of tillers initiated in a week
depends on the temperature of the previous week, and on a shoot production rate. Tiller production stops at
double ridge stage. The survival of each tiller group is calculated from the accumulated thermal time since double
ridge, taking into account the existing shoot density at the birth of each tiller group. Higher is this density, more
reduced is die proportion of surviving tillers. In a practical way. it means that the first bom tillers have a chance to
survive of about 0.9 or 0.95, when the tillers bom near the double ridge stage generally do not survive. At
anthesis stage, tiller death stops, and all live shoots are assumed to cany ears.
Leaf growt h and senescence- The maximum dimensions of the leaves are defined and depends on the position of
the leaves an the stem. Each leaf reaches its final size in 1.8 phyllochrons. The model simulates the leaf growth
by describing the daily length and width growth of both laminae and sheaths. Because various observations
during the main period of vegetative growth showed that there are generally between 3 and 4 active green leaves
on a shoot, the thermal time required from the attainment of maximum size to zero active area is estimated to be
3.5 time the phyllochron interval far most of the leaves. Each leaf remains totally active before the b eginning of
the senescence during approximately the 2 thirds of that time.
23 Radiosity experiment
We want to examine the results of a radiosity computation at early stage of the growing season of the winter
wheat The program of the functional runs with a daily time step. When the day processed is a day chosen for the
computation of the radiosity, all the variables related to the description of die structure are written in an output
file. It means: a) per square meter main shoot population, population of the first category of tillers (on the first
leave of the main shoot), of the second category, etc b) dimension of all the existing leaves: width and length
of lamina and sheat areas, c) percent of senescent area on the various leaves.
To keep computation time (which is very hi gh with the radiosity technique) w ithin reasonable
limits, we restrict our study area to a quater of square meter. Then, with an original sowing density of 250 shoots/
square meter, we worked with 63 main shoots on our study area, which allows an easy representation in 7 rows of
9 shoots. The space between the rows is approximately 7 cm, and die space between 2 shoots in the same row is
between 5 and 6 cm. The x y positions of the main shoots are then defined. The x y position of any tiller is the x y
position the main shoot where this tiller appeared, plus a deviation of 1 or 2 cm following an «ximnth direction
which is randomly chosen.
Some other information, which are not simulated by the model are needed to use the radiosity
techniques: the height and the orientation of each element. Many graphic representations or pictures of a wheat
shoot at various phenological stages have been consulted in general agronomic literature (e.g. Soltner, 1978), as
well as some reported values of height or leaf mr.linatirm which were found in published studies. The values used
in that work are arbitrary, even if they are carefully chosen. In a future study, the use of empirical relation should
be used to add to the canopy development submodel, the simulation of the position (especially the height) and the
orientation of the leaves. For same particular days, we present now the structure of the canopy and our way to