440
tabliehen Stadium begrundigt sich auf Symptome in Einzel
pflanzen im dem Unterwuchs. Zuverlassliche Entdeckung der
Krankheitssymptome kann mann durch grossmasstäbliche Photo
graphien erreichen, um die Zuverlässlichkeit des kleinmass-
täblichen Stadiums zu verbessern bedarf es dagegen ferner
Forschung.
INTRODUCTION
The jarrah ( Eucalyptus marginata Sm) forests of
western Australia are severely affected by "jarrah dieback",
a disease caused by the root rotting pathogen Phytophthora
cinnamomi Rands (Podger, Doepel and Zentmyer 196> 5)• First
reported in 1921 in small areas of the northern jarrah
forest, the disease now assumes major proportions with an
estimated 170,000 ha of forest affected by the disease
(Forests Department 1974)• Reliable mapping of the disease
is necessary for overall resource planning, scheduling
salvage operations and for the forest hygiene program aimed
at reducing the spread of the disease (Batini and Hopkins
1 972).
The symptoms of the disease are decline and death
of the jarrah as well as a wide range of under storey and
shrub species (Podger 1972). The classical manifestation
of the disease is the yellowing and death of a number of
understorey and shrub species, the most obvious ones being
bull banksia ( Banksia grandis Willd.) . zamia palm ( Macrozamia
re idle i O.A. G-ardn.) and blackboy (Xa nthorrhoea preissii
Endl.) , tliough the latter may frequently survive even in
advanced dieback areas. Obvious deterioration of the jarrah
crowns is usually not apparent until some time after symptoms
appear in the understorey, but ultimately the jarrah becomes *
thin crowned, and the tree eventually dies. The smaller
and less obvious susceptible shrub species also die out
during this process leaving a stand of dead jarrah with a
light shrub cover.
Marri ( E. calophylla R.Br.) . the other main tree
species of the jarrah forest association is resistant to
P. cinnamomi and together with other resistant shrub species
eventually recolonises the site. A transect through a
classical dieback site exhibits all stages of the disease
from the first deaths of the banksia through to stands of
affected and dead jarrah with live marri. Variations from
this pattern are not uncommon due to varying degrees of sus
ceptibility, rate of fungal activity and to the variable
proportions of susceptible and resistant species in the
understorey. These variations sometimes cause difficulty
in deciding whether or not an area is affected by P.cinnamomi .
In the northern jarrah forest region (Figure 1),
the mapping of dieback from 1: 40,000 panchromatic aerial
photographs has been successful for detection of the disease
at about the stage where the jarrah itself is being affected.