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ACKNOWLEDGEMENTS 
The authors gratefully thank Dr. D.G. Rossiter for insightful 
comments and practical foundation for statistical software R, 
and Dr. F. Corsi for providing basis of ecological modelling 
with GIS. Funding for this research was provided by 
International Institute for Geo-information Science and Earth 
Observation (ITC), “Research promotion fund in remembrance 
of Mori Taikichiro in 2007” of Keio University and “Grant-in- 
Aid for Project by SFC Research Institute in 2007” of Keio 
University.