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must have migrated forwards and backwards, changing
their locality but not their climate, and remaining always
“unchanged by changing place.” If, then, a given species
could manage through the geological ages to find a sphere
similar to the one to which it had been long ago adapted,
there seems no reason why it should undergo modification,
at any rate through the influence of external conditions.
In the second place we learn from the experience of
artificial selection that, while conditions remain unchanged,
those variations which necessarily arise in connection with
sexual reproduction do not lead to the establishment
of new varieties, races, or species, apart from the principle
of selection. Mr. Darwin illustrates this law by comparing
the gold-fish and the carp. The former are kept in glass
vessels and have been made the subjects of artificial
selection and have therefore yielded many races. The
carp have remained unmodified, because artificial selection
could not be brought to bear upon them. These facts
illustrate the general principle that, without the isolation
of similar variants for breeding purposes, the variations
which are necessarily associated with sexual reproduction
will be swallowed up in the average of the race by virtue
of the principle of regression to mediocrity, and thus
the characteristic type of a species will be maintained.
It is further very important to observe in this connection
that a very great amount of individual variety is con
sistent with the continued stability of species. Nor does
the idea of the stability of species exclude the idea of
a severe struggle for existence ; for it is almost impossible
to conceive that the species which have undergone little or
no change through all the æons of geological time can
have been altogether free from the strife. And this
probability becomes a certainty when we consider what
actually takes place at the present time. When circum-