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whole adaptive morphology of nature. How immense, thei'efore,
must be the number of unuseful variations. These are probably
many hundred of times more numerous than the useful variations,
although they are all, as it were, stillborn or allowed to die out imme
diately by intercrossing.”—{Journal 'of Linnean Society. Zoology,
vol. xix., p. 348.)
If this view is the correct one, Natural Selection secures
the survival of a few favourable variations and the des
truction of a few unfavourable ones, but does not prevent
a large number of unuseful variations from surviving;
but it is assumed that these will be stillborn or die out
immediately by intercrossing. Here, then, the principle of
regression to mediocrity does more than Natural Selection
to maintain the efficient stability of species.
But it is not at all so clear that these unuseful varia
tions will be stillborn and will be immediately absorbed
in the average of the race ; and, if not, there is no reason
why they should not intermarry with the more typically
perfect ; and in this way the average—not the ideal
perfection—of the race would be maintained.
If, now, we take the hypothesis that the variants range
themselves in a central column, on either side of which
there are struggling variants, then the principle of regres
sion to mediocrity will do all that is required ; the plus
and the minus will neutralise one another, and Natural
Selection is not required.
Next, let us suppose that the variants range themselves
in two columns on either side of a central line. In the
first place, we must be well assured that these variants
are real sheep and goats—all useful on the one side and
all prejudicial on the other side. Only on this hypothesis
could Natural Selection act in the way supposed ; for if
there were a large output of unuseful variations, Natural
Selection could not touch them. But if Natural Selec
tion did choose all the column to the right because it