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past, has most likely been the active origin of the
amblystoma race from the axolotl stock. Presumably
the axolotl as the gill-bearing form is the primitive stock,
the amblystomas being a derived race, but nevertheless
representing a true species, of which the axolotl may be
regarded as the larval form. The shrivelling of the gills,
seen in the experiments on the axolotl, was probably due
to a mechanical cause—that of dryness of the surroundings.
Dr. Weismann, who believes that the axolotl is a de
generate form, declares that the transformation of the
axolotl into the amblystoma was not brought about by
Natural Selection.
“ If their gills by this means become shrivelled up or completely
disappear, even this is not adaptation in the Darwinian sense, but the
effect of directly-acting external influences and chiefly of diminished
use.”—(Studies in the Theory of Descent, ft. jgi.)
Mr. W. Mattieu Williams, speaking of the transforma
tion of the Mexican axolotl into an amblystoma, says:—
“ Here is a case of very rapid modification, in which the animal,
without the aid of hereditary variation, without any destruction of the
unfit and survival of the fittest, has at once adapted itself to varying
external conditions—a case apparently supporting the old-fashioned
development theories of the predecessors of Darwin.”—{Gentleman's
Magazine, vol. cclxii., ft. 100.)
(d) PALEONTOLOGY.
“ Bones bear witness.”
—Comedy of Errors. Act iv., sc. 4.
It is the opinion of some writers that palaeontology does
not afford any conclusive evidence of the action of Natural
Selection.
“ While the general testimony of palaeontology is overwhelmingly
in favour of the view that some general law of evolution has operated