One spectral components analysis
equation (Wiegand and Richardson, 1984)
is
FPAR(VI) = FPAR(L) x L(VI) [1]
read as FPAR as a function of any
vegetation index (VI) dominated by the
NIR reflectance of the canopy equals
FPAR as a function of leaf area index
(L) times L as a function of VI. The
equation states that because there is a
functional relation between FPAR and L
and between L and VI it follows that
there is also one between FPAR and VI.
Calibration of FPAR directly in terms
of VI avoids the tedious labor of
determining L, but more importantly,
makes FPAR estimates available for many
remotely observable fields. It is now
recognised that FPAR is a nearly linear
function of VI (e.g. Gallo et al.,
1985; Wiegand and Richardson, 1987),
and attempts at the biophysical
explanation of the linearity have been
made (Sellers, 1985, 1987; Choudhury,
1987).
Fig. 1 illustrates Eq. [1]
relationships for three different
crops: cotton (Gossypium hirsutum, L.),
wheat (Triticum aestivum, L.), and corn
(maise) (Zea mays, L.). The equations
that express the 18 functional
relations of Fig. 1 are summarized in
Table 4 of Wiegand and Richardson
(1990b.)
Remote observations of the canopies,
expressed in Fig. 1 as PVI, are the
independent variable used to estimate L
in the second right side term of Eq.
[1]. That L can be estimated from
spectral vegetation indices, and that
the fraction of the PAR that is
absorbed can be estimated from L are
well accepted. The functional
relations of the two right side terms
are both nonlinear while the left side
functional relation is linear (Fig. 1).
The significance of Eq. [1] is that
FPAR can be estimated from remote
observations. Since APAR drives
photosynthesis, the process by which
plants produce the assimilates for
growth and reproduction, VI provide a
way to monitor plant development and
yield. The linearity of the relation
makes it easy to use.
Economic Yield from One or a Few
Observations per Season
An equation that contains the L(VI)
term of Eq. [1] and incorporates
economic yield (Y) is
Y(VI) = L(VI) x Y(L) [2]
where Y is the salable plant part as
appropriate for the crop (grain, root,
fruit, fiber, or aboveground biomass),
g/m 2 . Eq. [2] relates the
photosynthetic capacity of the crop
characterized by L and VI to its
economic yield.
Eq. [2] applies when the number of
remote observations is limited to one
or a few per season between late
vegetative and mid-reproductive
stages. Observations during this
interval are optimal because
measurements of VI too early in the
season or too late into senescence are
not representative of the canopies'
photosynthetic size to support creation
and growth of the plant parts that
constitute yield. For cereals such as
maize, sorghum, rice, and wheat, this
optimal period lasts from late stem
extension to about the milk stage of
the grain. During this interval L is
at a stable, plateau value. Other
crops as diverse as cotton, potato, and
various melons also reach a plateau
value of L during reproduction because
most of the assimilates of
photosynthesis is translocated to and
stored in the reproductive organs; new
growth of foliage is slow and part of
it replaces leaves that senesce.
Fig. 2 illustrates Eq. [2]
relationships for rice (Oryza sativa,
L.) adapted from Shibayama et al.
(1988) and Wiegand et al. (1989). The
data are from 13 treatments that
consisted of imcomplete combinations of
3 cultivars, 2 transplanting dates
three weeks apart, and 6 nitrogen
application rates. In Fig. 2, part 'c'
depicts the second right side term,
part ’b' the first right side term, and
part 'a' the left side term of Eq. [2],
respectively. The Lh and PVIh in Fig.
2 designate averages of these variables
by treatment for three dates
surrounding heading.
As was characteristic of Eq. [1], the
right side terms in Eq. [2] depicted in
Fig. 2 are nonlinear whereas the left
side term is linear. Diversity in the
treatments pooled for part 'c' and
experimental error in all measurements
account for the scatter in part 'a'.
In spite of the data limitations, 63%
of the yield variation is attributable
to PVIh and the relation is
unmistakeably linear.
Fig. 3 illustrates Eq. [2]
relationships for grain sorghum
(Sorghum bicolor L. Moench) after
Wiegand and Richardson (1984). Leaf
area index was measured periodically
during the growing seasons of 1973,
1975, and 1976 in farmers' fields,
Landsat MSS digital counts acquired
during grain filling were extracted
from computer compatible tapes provided
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